Notes – Seed: Structure, Types & Germination
Life
of a Plant Starts with a Seed:
- Every plant begins its life as a
seed.
- A seed is crucial for the
development of a new plant.
Seed
Formation:
- A seed develops from an ovule.
- This happens after fertilization
(the fusion of male and female gametes).
Dormancy
in Seeds:
- Under unfavourable conditions
(like lack of water or suitable temperature), seeds enter a dormant state.
- In dormancy, the seed is inactive
and does not germinate.
Germination
in Favourable Conditions:
- When favourable conditions
(like proper moisture, warmth, and air) are available, the seed begins to
germinate.
- Germination is the process where
the seed starts to grow into a new plant.
Food
Storage in Seeds:
- During the initial stages of
germination, the seed uses the food stored within it to grow.
- This stored food helps the
seedling develop until it can perform photosynthesis.
Definition
of Seed:
- A seed is the ripened ovule of a
plant.
- It contains an embryo that
germinates into a new plant.
Parts
of a Seed:
- A mature seed consists of two
essential parts:
- Seed coat
- Embryo
Seed
Coat:
- A protective outer covering of
the seed, developed from the ovule's integuments.
- The seed coat has two layers:
- Testa: Outer layer
- Tegmen: Inner layer
- Hilum: A scar on the seed, marking the attachment point to its
stalk.
- Micropyle: A small opening near the hilum, helps in absorbing water
during germination and allows the diffusion of respiratory gases.
Embryo:
- The young or miniature plant
enclosed within the seed coat.
- The embryo consists of five
distinct parts:
- Cotyledons: Seed leaves; two in dicots, one in monocots.
- Plumule: Gives rise to the shoot upon germination.
- Epicotyl: The part of the embryonic axis above the cotyledon
attachment, holds the plumule.
- Hypocotyl: The part of the embryonic axis below the cotyledon
attachment.
- Radicle: Located at the tip of the hypocotyl, develops into the
root during germination.
Dicotyledonous
Seeds:
- Have two cotyledons.
- Can be either exalbuminous
(without stored food, e.g., pea, gram, bean) or albuminous (with
stored food, e.g., castor, cotton).
Monocotyledonous
Seeds:
- Have one cotyledon.
- Can be either exalbuminous
(e.g., Alisma) or albuminous (e.g., maize, wheat, rice).
Germination
Process:
- Radicle: Develops into the root.
- Plumule: Develops into the shoot.
Food
Storage in Seeds
- Food Storage Locations:
- In seeds,
food is stored in two main places:
- Cotyledons: The first leaves of the seed that store food.
- Endosperm: A special tissue that also stores food.
- Types of Seeds Based on Food
Storage:
- Non-Endospermic
(Exalbuminous) Seeds:
- Example
seeds: Gram, Pea, Bean, Mustard.
- Food is
stored in cotyledons.
- Endosperm is absent, indicating that it has been fully used by the
developing embryo.
- Endospermic
(Albuminous) Seeds:
- Example
seeds: Castor, Maize, Rice.
- Food is
primarily stored in the endosperm.
- These
seeds contain a significant amount of stored food for the embryo's
development.
- Structure of Fruits and Seeds:
- Fruit:
- It is the
enlarged, ripened ovary of a flower.
- The outer
layer of the ovary forms the fruit wall, which encloses the seed.
- The fruit
protects the seed and aids in its dispersal.
- Seed:
- It is the
ripened ovule that contains the embryo, which will develop into a
future plant.
- The seed
coat serves as a protective layer for the embryo.
- Grain:
- It is a fruit
where the fruit wall and the seed coat are fused together,
creating a protective layer around the seed.
Structure
of Dicotyledonous Bean Seed
- Shape and Covering:
- The bean
seed is kidney-shaped.
- It is
covered by a hard seed coat.
- Seed Coat Layers:
- The seed
coat has two layers:
- Testa: The outer layer of the seed coat.
- Tegmen: The inner layer of the seed coat.
- These
layers are fused together to provide protection to the
embryo.
- Hilum:
- Located on
the concave side of the seed.
- It is a whitish
oval scar where the seed was attached to the pod via a structure
called the funicle.
- Micropyle:
- A small
opening located close to the hilum.
- It allows
the seed to absorb water during germination.
- It also
serves as a passage for the diffusion of respiratory gases.
- Raphe:
- An elongated
ridge found on the testa at the other end of the hilum.
- Inner Structure:
- When the
seed coat is peeled off, a white fleshy body is revealed,
consisting of:
- Two fleshy
cotyledons: These store food for
the plant.
- Short axis: To which the cotyledons are attached.
- Axis Components:
- The axis
consists of:
- Radicle: Located outside the cotyledons; it develops into the root
system.
- Plumule: Develops into the shoot system.
- The axis is
divided into:
- Hypocotyl: The part below the cotyledon attachment.
- Epicotyl: The part above the cotyledon attachment.
Structure
of Monocotyledonous Maize Grain (Zea mays)
- Type of Fruit:
- Maize is a one-seeded
fruit known as a caryopsis.
- The fruit
wall (pericarp) is fused with the seed coat.
- External Appearance:
- The maize
grain has a flat, almost triangular, and oblong
shape.
- It has a large
yellowish upper area indicating the position of the endosperm.
- The lower
side contains a small whitish area which houses the embryo.
- Seed Coat:
- The seed
coat is not distinct as it is fused with the fruit wall
(pericarp).
- Micropyle and hilum are also not distinguishable.
- Internal Structure:
- In a longitudinal
section, it reveals two distinct regions:
§ Upper
region: Endosperm
§ Lower
region: Embryo
- Endosperm:
- The
endosperm is surrounded by a one-cell thick layer called the aleurone
layer.
- It contains
aleurone grains, which are proteinaceous in nature.
- Embryo Structure:
- The embryo
consists of:
§ Single
Cotyledon called scutellum: it is shield-shaped &
flat, closely pressed against the endosperm.
§ Function:
Absorbs food from the endosperm and transfers it to the growing parts of the
embryo.
§ Plumule:
The growing tip of the shoot, accompanied by a few embryonic leaves.
§ It
is covered by a sheath called coleoptile.
§ Radicle:
Located at the base of the grain and covered with a sheath called coleorrhiza.
§ Hypocotyl:
Very short and represented by a short axis between the radicle and plumule.
Seed
Germination
- Seed Condition at Shedding:
- Most seeds
are shed from the parent plant in a dry condition.
- In this
state, physiological activities are very slow, and minimal food is
utilized.
- Respiration in seeds is also slow during this dry phase.
- Dormancy:
- Seeds can
remain alive and dormant for extended periods when dry.
- Some seeds
can germinate only a few days after being shed, after which they lose the
ability to germinate.
- Dormant
seeds do not germinate even when provided
with all necessary conditions for germination. This condition is known as
dormancy.
- Germination Process:
- Germination
is the process where the food reserves in a seed are broken down,
allowing the embryo to start growing.
- It begins
when the seed receives favourable conditions (e.g., water, warmth,
and oxygen).
- Irreversibility of Germination:
- The process
of germination is irreversible.
- Once
germination starts, it cannot stop, and the seed cannot return to the
dormant state.
Key
Terms:
- Dormancy: A state in which seeds remain alive but do not germinate.
- Germination: The process of growth from a seed into a new plant.
Conditions
Necessary for Seed Germination
- Water
- Rapid
Uptake: Germination begins with the absorption
of water through the micropyle and seed surface.
- Functions
of Water:
- Softens
the seed coat, allowing it to rupture for plumule (shoot) and radicle
(root) emergence.
- Accelerates
metabolic activities within the seed.
- Aids in
hydrolysis, converting stored food into soluble products.
- Facilitates
translocation of soluble nutrients from storage tissues to the
developing embryo.
- Hastening
Germination:
- Soaking
seeds in water before sowing.
- Removing
the seed coat.
- Availability of Oxygen
- Energy
Requirement: The embryo needs energy to
resume growth during germination.
- Source of
Energy: Energy is obtained through the
oxidation of food stored in cotyledons or endosperm, which requires
oxygen.
- Favourable Temperature
- Physiological
Processes: Various physiological
processes take place during germination; therefore, a suitable
temperature is essential.
- Optimum
Temperature Range:
- Varies by
seed type, but most seeds do not germinate below 0°C or above 48°C.
- Ideal
temperature range for most seeds: 25°C to 35°C.
- Light Conditions
- Some seeds
require light for germination, while others germinate only in darkness.
The
Germination Process
- Definition: Germination is the process by which a seed develops into a new plant.
- Initiation:
- Germination
begins with the rapid uptake of water by the seed through a small
opening called the micropyle.
- This water absorption causes the seed to swell, resulting in an increase in weight.
- Physical Changes:
- The seed
coat becomes soft due to the absorption of water.
- The first visible sign of germination is the swelling of the seed.
- Physiological Changes:
- The absorption
of water triggers several physiological changes in the seed.
- There is an increase in respiratory activity in the germinating seeds.
- Enzyme Production:
- The embryo
inside the seed produces enzymes that convert stored food
materials (found in cotyledons or endosperm) into soluble
forms.
- These soluble forms are essential for the growing embryo's nutrition.
- Cell Division:
- Once food
becomes available, cell division begins in the growing embryo,
particularly in two parts:
- Radicle (the embryonic root)
- Plumule (the embryonic shoot)
- Emergence of Structures:
- The growth
of the embryo tissues causes the seed coat to rupture.
- The radicle
is the first part to emerge from the seed coat, forming the root
system, which grows downwards into the soil.
- After the
radicle, the plumule emerges and develops into the shoot of
the plant.
Result
of Experiments on seed Germination: (Refer to your book for the experiments).
- Result
(Experiment 1) - Pyrogallic acid absorbs oxygen from the air. Without Oxygen,
seeds do not germinate in Flask A.
- Result (Experiment 2) - Adequate water
must be present for germination to start and progress. Excess water prevents
germination, probably by excluding oxygen.
- Result
(Experiment 3) – Suitable temperature is necessary for germination.
Three
bean seeds experiment (Refer to your book for the experiment)
- The top seed does not germinate
because it gets only oxygen and no water.
- The bottom seed receives water but
very little oxygen.
- The middle seed germinates properly as
it receives both oxygen and water.
Types
of Seed Germination
-
Seed germination is the process by
which a seed develops into a new plant.
-
The behaviour of the cotyledons (the
first leaves) during germination distinguishes two main types of seed
germination:
1.
Epigeal Germination
2.
Hypogeal Germination
Epigeal
Germination
- Definition: A type of germination where the cotyledons are lifted above
the ground.
- Mechanism:
- Occurs due
to the rapid elongation of the hypocotyl (the part of the embryonic axis
just below the cotyledons).
- Examples: Seen in seeds such as:
- Kidney bean,
Castor, Cotton, Sunflower, Tamarind
- Root System:
- The primary
becomes the main root.
- Outcome: The cotyledons are carried above the soil surface during
germination.
Hypogeal
Germination
- Definition: A type of germination where the epicotyl elongates quickly.
- Mechanism:
- The epicotyl
(the part of the embryonic axis above the cotyledons) pushes the plumule
(young shoot) up into the air.
- The cotyledons
remain on or below the ground.
- Examples: Observed in seeds like:
- Gram, Pea, Mango,
Maize
- Root System:
- The primary
root does not become the main root.
- Adventitious
roots develop close to the primary root, forming a fibrous root system.
Viviparous
Germination
- Definition: A special type of germination where seeds germinate while
still attached to the parent plant.
- Example: Common in mangrove plants, such as:
- Rhizophora,
Sonneratia
EXERCISES
1. Multiple Choice Questions (MCQs)
(i) How is a seed formed?
Answer: (a) It develops from an ovule after
fertilisation.
(ii) A mature seed consists of the following
parts:
Answer: (c) Seed coat and embryo
(iii) The embryo of a mature seed consists of:
Answer: (d) Cotyledons, plumule, epicotyl,
hypocotyl, and radicle.
(iv) Cotyledons are also known as:
Answer: (a) Seed leaves.
(v) The fruit called caryopsis is found in:
Answer: (c) Maize.
(vi) Aleurone layer present in maize seed is
rich in:
Answer: (c) proteins.
(vii) Epigeal germination is seen in which of
the following seeds?
Answer: (b) Gram.
(viii) Which type of germination takes place
in mango?
Answer: (a) Hypogeal germination.
(ix) Which of the conditions are essential for
seed germination?
Answer: (d) All of these (Water, Oxygen,
Optimum temperature).
(x) The first visible indication of
germination is:
Answer: (a) The swelling of the seed.
(xi) Assertion (A): The bean seed is
kidney-shaped and remains covered by a hard seed coat. Reason (R): Bean seed is
dicotyledonous.
Answer: (a) Both A and R are true.
(xii) A bean seed is placed in a moist paper
towel and kept in a warm, dark place. A few days later, the seed germinates,
and a small plant begins to grow. Which of the following is/are involved in the
process described above?
Answer: (e) All the above
2.
Definitions
(i) Embryo:
The embryo is the young plant that
develops from the fertilized ovule and consists of the radicle, cotyledons, and
plumule.
(ii) Exalbuminous seed: An
exalbuminous seed is a seed that does not contain endosperm. The cotyledons
store the food and are fleshy.
(iii) Germination: Germination
is the process by which a seed develops into a new plant when the conditions
are favourable.
(iv) Seed dormancy: Seed dormancy is a period during which seeds
remain alive, but do not germinate, even under favourable conditions.
3. Functions of the Following Parts
(i) Coleoptile: The
coleoptile is the protective sheath covering the emerging shoot of a seedling
and helps it grow upwards through the soil.
(ii) Coleorrhiza: The coleorrhiza is the protective sheath
surrounding the radicle in monocot seeds, aiding in its emergence.
(iii) Aleurone layer: The aleurone layer stores proteins and enzymes
and plays a crucial role in the early stages of germination by providing
nutrients.
(iv) Micropyle: The micropyle is a small opening in the seed
coat through which water and oxygen can enter, facilitating the germination
process.
(v) Endosperm: The endosperm provides nutritional support to
the developing embryo, supplying it with the necessary energy for growth.
(vi) Plumule: The plumule is the part of the embryo that
develops into the shoot and leaves of the new plant.
4,
Differentiate between the following:
(i)
Epicotyl and hypocotyl
Question |
Epicotyl |
Hypocotyl |
Definition |
The part of the
seedling above the cotyledons. |
The part of the
seedling below the cotyledons. |
Location |
Located above the
cotyledons. |
Located below the
cotyledons & above the radicle. |
Function |
Develops into the stem &
leaves of the plant. |
Supports the plant &
connects the radicle to the epicotyl. |
Appearance |
Typically elongated &
greenish in color. |
Generally short &
cylindrical. |
(i) Radicle and plumule
Question |
Radicle |
Plumule |
Definition |
The
embryonic root of the seed. |
The
embryonic shoot of the seed. |
Location |
Located
at the lower end of the seed. |
Located
at the upper end of the seed. |
Function |
Develops
into the primary root of the plant. |
Develops
into the stem and leaves of the plant. |
Appearance |
Generally
straight and may be fleshy. |
Generally
short and may have leaf-like structures. |
(iii) Bean Seed and Maize seed.
Feature |
Kidney Bean Seed |
Maize Grain |
1.
Seed Coat |
-
Thin and brownish in colour. |
-
Very thin and fused with the pericarp. |
2.
Hilum & Micropyle |
-
Both are visible. |
-
Both are not visible. |
3.
Cotyledons |
-
Contains two thick fleshy cotyledons. |
-
Contains a single cotyledon. |
4.
Endosperm |
-
Absent in the seed. |
-
Present as a part of the seed. |
5 Answer the
following:
(i). Describe the
structure of a seed.
A seed typically
consists of three main parts:
- Seed Coat: The outer protective layer that shields the seed
from damage and prevents water loss.
- Cotyledons: These are the seed
leaves. In dicot seeds, there are usually two cotyledons, while monocot seeds
have one. Cotyledons provide nourishment to the developing plant during
germination.
- Embryo: The part of the seed that develops into the new
plant. It contains the embryonic root (radicle), which will grow into the root,
and the embryonic shoot (plumule), which will develop into the stem and leaves.
(ii). What conditions
are essential for seed germination? Describe their importance.
The essential
conditions for seed germination include:
- Water: It is necessary for the seed to swell and soften the
seed coat. Water activates enzymes that begin the metabolic processes necessary
for growth.
- Oxygen: Seeds require oxygen for respiration. This process
provides energy needed for growth and development.
- Temperature: Most seeds require a
specific temperature range (usually warm) to germinate effectively. Optimal
temperatures enhance enzyme activity and metabolic processes.
- Light (for some seeds): Certain seeds need light to
germinate, while others do not. Light can trigger specific germination
processes in some species.
(iii). Differentiate
between hypogeal and epigeal germination.
Aspect |
Epigeal Germination |
Hypogeal Germination |
Cotyledon Position |
Cotyledons are carried
up and out of the soil. |
Cotyledons remain below
ground. |
Growth Mechanism |
Caused by the growth of
the hypocotyl |
Caused by the growth of
the epicotyl |
Plumule Position |
The plumule tip lies
between the cotyledons. |
The plumule is carried
up in the air above the cotyledons. |
Kidney bean, castor, cotton, sunflower, tamarind, etc. |
Gram, pea, mango, maize, etc. |
(iv). Give two
examples each of the following types of seeds:
(i) Dicotyledonous
albuminous seed: Castor bean, Sunflower seed.
(ii) Dicotyledonous
exalbuminous seed: Pea seed, Bean seed.
(iii) Monocotyledonous
albuminous seed: Maize grain, Rice grain.
(v). How would you
demonstrate that oxygen and water are necessary for seeds to germinate?
We can conduct a
simple experiment:
1. For Water:
- Let us take two identical seeds (e.g.,
beans) and place one in moist soil and the other in dry soil.
- Observation over a few days - The seed in
moist soil will germinate, while the one in dry soil will not.
2. For Oxygen:
- Let us take two containers with moist soil
and place a seed in each.
- Let us seal one container with a lid
(preventing oxygen entry) and leave the other open.
- Observation over a week - The seed in the
open container will germinate, while the sealed one will not.
(vi). Why does a
farmer plough his field and make the soil loose before sowing seeds?
A farmer ploughs the
field to:
·
Improve Soil Aeration: Loose soil allows
better air circulation, which is essential for root growth and helps in oxygen
availability for germinating seeds.
·
Enhance Water Absorption: Loosened soil absorbs
water more effectively, ensuring that seeds have adequate moisture for
germination.
·
Reduce Soil Compaction: Ploughing breaks up
hard soil, preventing the roots from having difficulty penetrating the ground.
6. Draw labelled
diagrams showing the structure of the following seeds:
(i) Kidney
Bean - Do it Yourself
(ii) Maize
Grain - Do it Yourself
7. Outline the stages
in the germination of the following seeds with the help of labelled diagrams:
(i) Kidney
Bean Seed - Do it Yourself
(ii) Maize
Grain - Do it Yourself
8. Observe the figure given below and answer the following:
(i) Label the parts
marked 1-5.
1
– Cotyledon
2
– Plumule
3
– Epicotyl
4
– Hypocotyl
5 – Radicle
(ii) Identify whether
the given seed is a monocot seed or a dicot one. Give one reason in support of your
answer.
The given seed is a dicot, as it contains two
cotyledons.
9. Observe the given figure and answer the following:
(i) Identify the given seed as monocot
or dicot.
Monocot
(ii) Give one reason in support of your
answer.
It
contains only one cotyledon
(iii) Label the parts marked 1-7.
1 – Pericarp and seed
coat fused
2 – Endosperm
3 – Coleoptile
4 – Plumule
5 – Cotyledon (Scutellum)
6 – Radicle
7 – Coleorrhiza
10. Observe the figure given below and answer the following:
(i) What does the given figure show?
Epigeal
Germination
(ii) Label the parts marked 1-5.
1 – Radicle
2 – Cotyledon
3 – Hypocotyl
4 – Cotyledon
5 – Foliage Leaf
(iii) Name one plant in which this
takes place.
Kidney bean
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